Papers available on request, if not open access.

Book chapters

  1. de los Santos CB, Scott A, Arias-Ortiz A, Jones B, Kennedy H, Mazarrasa I, McKenzie L, Nordlund LM, de la Torre-Castro M. 2020. Seagrass ecosystem services: Assessment and scale of benefits. In: Out of the blue: The value of seagrasses to the environment and to people, edited by Potouroglou M, Grimsditch G, Weatherdon L, Lutz S. Nairobi, Kenya. United Nations Environment.

Scientific articles

  1. Cozzolino L, de los Santos CB, Zardi G, Repetto L, Nicastro K. Accepted. Microplastics in commercial bivalves harvested from intertidal seagrasses and sandbanks in the Ria Formosa lagoon, Portugal. Marine and Freshwater Research.
  2. de los Santos CB, Krång A-S, Infantes E. 2021. Microplastic retention by marine vegetated canopies: simulations with seagrass meadows in a hydraulic flume. Environmental Pollution, 269: 116050 [Open Access].
  3. Mishra AK, Cabaço S, de los Santos CB, Apostolaki ET, Vizzini S, Santos R. In press. Long-term effect of elevated CO2 on the population dynamics of the seagrass Cymodocea nodosa: evidence from volcanic seeps. Marine Pollution Bulletin.
  4. de los Santos CB, Olive I, Moreira M, Silva A, Freitas C, Araújo Luna R, Quental-Ferreira H, Martins M, Costa MM, Silva J, Cunha ME, Soares F, Pousão-Ferreira P, Santos R. 2020. Seagrass meadows improve inflowing water quality in aquaculture ponds. Aquaculture 528, 735502.
  5. Cozzolino L, Nicastro K, Zardi G, de los Santos CB. 2020. Species-specific plastic accumulation in the sediment and canopy of coastal vegetated habitats. Science of the Total Environment, 723: 138018.
  6. Franco JN, Arenas F, Sousa-Pinto I, de los Santos CB. 2020. Snapshot of macroalgae and fish assemblages in temperate reefs in the Southern European Atlantic ecoregion. Diversity, 12(1): 26. [Open Access]
  7. Barracosa H, de los Santos CB, Martins M, Freitas C, Santos R. 2019. Ocean literacy to mainstream ecosystem services concept in formal and informal education: the example of coastal ecosystems of Southern Portugal. Frontiers in Marine Sciences, 6: 626. [Open Access]
  8. Paul M de los Santos CB. 2019Variation in flexural, morphological and biochemical leaf properties of eelgrass (Zostera marina) along the European Atlantic climate regions. Marine Biology, 166:127. Equal contribution.
  9. de los Santos CB, Krause-Jensen D, Alcoverro T, Marbà N, Duarte, CM, van Katwijt MM, and 21 more authors. 2019Recent trend reversal for declining European seagrass meadows. Nature Communications, 10: 3356. [Open Access]
  10. de los Santos CB, Arenas F, Neuparth T, Santos MM. 2019. Interaction of copper pollution and ocean acidification in seagrass ecosystems: toxicity, bioconcentration and dietary transfer. Marine Pollution Bulletin, 142: 155-163.
  11. Santos R, Duque-Núñez N, de los Santos CB, Martins M, Carrasco AR, Veiga-Pires C. 2019. Superficial sedimentary stocks and sources of carbon and nitrogen in coastal vegetated assemblages along a flow gradient. Scientific Reports, 9: 610. [Open Access]
  12. de los Santos CB, Godbold JA, Solan M. 2017. Short-term growth and biomechanical responses of the temperate seagrass Cymodocea nodosa to CO2 enrichmentMarine Ecology Progress Series, 572: 91-102. [Open Access]
  13. de los Santos CB, Onoda Y, Vergara JJ, Pérez-Lloréns JL, Bouma TJ, La Nafie YA, Cambridge ML, Brun FG. 2016. A comprehensive analysis of mechanical and morphological traits in temperate and tropical seagrass species. Marine Ecology Progress Series, 551: 84-91.
  14. Santos MM, Ruivo R, Marques M, Torres T, de los Santos CB, Castro F, Neuparth T. 2016Statins: An undesirable class of aquatic contaminants? Aquatic Toxicology, 174: 1–9.
  15. de los Santos CB, Vicencio-Rammsy B, Lepoint G, Remy F, Bouma TJ, Gobert S. 2016. Ontogenic variation and effect of collection procedure on leaf biomechanical properties of Mediterranean seagrass Posidonia oceanica (L.) Delile. Marine Ecology, 37: 750–759.
  16. de los Santos CB, Neuparth T, Torres T, Martins I, Cunha I, Sheahan D, McGowan T, Santos MM. 2015. Ecological modelling and toxicity data coupled to assess population recovery of marine amphipod Gammarus locusta: application to disturbance by chronic exposure to aniline. Aquatic Toxicology, 163: 60–70.
  17. Neuparth T, Martins C, de los Santos CB, Costa MH, Martins I, Costa PM, Santos MM. 2014. Hypocholesterolaemic pharmaceutical simvastatin disrupts reproduction and population growth of the amphipod Gammarus locusta at the ng/l range. Aquatic Toxicology, 155: 337–347.
  18. La Nafie YA, de los Santos CB, Brun FG, Supriadi M, van Katwijk MM, Bouma TJ. 2013. Biomechanical response of two fast-growing tropical seagrass species subjected to in situ shading and sediment fertilization. Journal of Experimental Marine Biology and Ecology, 446: 186–193.
  19. de los Santos CB, Brun FG, Vergara JJ, Pérez-Lloréns JL. 2013. New aspect in seagrass acclimation: leaf mechanical properties varies spatially and seasonally in the temperate species Cymodocea nodosa Ucria (Ascherson). Marine Biology. 160, 1083–1093. Erratum.
  20. La Nafie YA, de los Santos CB, Brun FG, van Katwijk MM, Bouma TJ. 2012. Waves and high nutrient loads jointly decrease survival and separately affect 2 morphological and biomechanical properties in the seagrass Zostera noltii. Limnology and Oceanography, 57: 1664–1672.
  21. de los Santos CB, Brun FG, Onoda Y, Cambridge ML, Bouma TJ, Vergara JJ, Pérez-Lloréns JL. 2012. Leaf-fracture properties correlated with nutritional traits in nine Australian seagrass species: potential implications for susceptibility to herbivory. Marine Ecology Progress Series, 458: 89–102. [Open Access]
  22. de los Santos CB, Brun FG, Bouma TJ, Vergara JJ, Pérez-Lloréns JL. 2010. Acclimation of seagrass Zostera noltii to co-occurring hydrodynamic and light stressesMarine Ecology Progress Series, 398: 127–135. [Open Access]
  23. Mercado JM,  de los Santos CB, Pérez-Lloréns JL, Vergara JJ. 2009Carbon isotopic fractionation in macroalgae from Cádiz Bay (southern Spain). Comparison with other biogeographic regions. Estuarine, Coastal and Shelf Science, 85: 449-458.
  24. de los Santos CB, Pérez-Lloréns JL, Vergara JJ. 2009. Photosynthesis and growth in macroalgae: linking functional-form and power scaling approaches. Marine Ecology Progress Series, 377: 113–122. [Open Access]


  • Martins M, de los Santos CB, Masque P, Carrasco AR, Veiga-Pires C, Santos R. Carbon and nitrogen stocks and burial rates in intertidal vegetated habitats of a mesotidal coastal lagoon.